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In many species, a prominent feature of sexual differentiation is the presence of a sexually dimorphic nucleus SDN in the preoptic hypothalamus, which is larger in males than in females.

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Most of the data suggests that homosexual rams, like female-oriented rams, are masculinized and defeminized with respect to mounting, receptivity, and gonadotrophin secretion, but are not defeminized for sexual partner preferences, also suggesting that such behaviors may be programmed differently. The heterosexual advantage hypothesis was given strong support by the Italian study demonstrating increased fecundity in the female matrilineal relatives of gay men.